The Fifty-Eighth Supplement to the AOU Check-list

Northern Shrike

It’s Christmas in July for most birders with the appearance of the now-annual Supplement to the AOU Check-list. This year, as always, Santa Claus giveth and Santa Claus taketh away. On balance, those who care about numbers will find their lists increasing. For the rest of us — for most of us — the yearly update is a chance to look into the workings of taxonomists and ornithologists as they toil to decipher the relationships among our birds.

thayer's gull 6

The greatest loss for listers is certainly that handsome gull “kind” known over the past 45 years as the Thayer gull. Jon Dunn and Van Remsen argued cogently, even devastatingly, that the research supporting full species status for the bird was thoroughly flawed, and that the “burden of proof” should be on those asserting its distinctness from the Iceland gull. To my memory, Dunn and Remsen’s is the only taxonomic proposal ever considered by the AOS committee to use the phrases “scientific misconduct.” The authors encourage further research into the taxonomy of the large herring-like gulls, but meanwhile, thayeri is reduced to a mere synonym. 

Eastern Willet

Some birders will probably be disappointed, too, by the committee’s having declined to accept a number of proposed splits and re-splits, some involving some of the most familiar birds on the continent. The willet remains a single species, as does the yellow-rumped warbler.

Myrtle Warbler


The eastern and western populations of the brown creeper, the Nashville warbler, and the Bell vireo were also sentenced to continued cohabitation.

But there are splits aplenty, too.

Baird's junco

The gorgeous little Baird junco gets its own box on the ticklist again, and the Talamanca hummingbird of Costa Rica and Panama is once again treated as distinct from the northerly Rivoli hummingbird.

magnificent hummingbird

To my surprise, we also have a new crossbill species in North America. The Cassia crossbill (the English name commemorates the type locality, and is far better than the cutesy scientific name sinesciuris) breeds in the South Hills and Albion Mountains of Idaho. It is apparently sedentary, making identification perhaps a bit easier; the bird is said to be larger than other sympatric crossbills, and to have different calls and songs.

My surprise has nothing to do with the quality of the research establishing this as a distinct species: all this genetics stuff is way beyond me. But I did not expect any real movement in crossbill classification to be inspired by one taxon; I’d thought the committee might wait for a universal solution to these difficult problems. In any case, Burley had better be ready for an ornitho-influx.

great gray shrike

We also get a split in the “gray” shrike complex. The North American northern shrike is now considered specifically distinct from its Old World counterparts; its species epithet is once again borealis, the name given it by Vieillot in 1808.

Northern Harrier

Our northern harrier is also split from the hen harrier of Europe, under the Linnaean name Circus hudsonius. The name honors the employer of James Isham, who sent the first specimens to George Edwards in the 1740s.

Common Redpoll darkish

The number of birders dreading the lump of the redpolls was almost as great as that of those devoutly wishing its consummation. The resolution (for now) leaves us with three species in the United States and Canada, the hoary, common, and lesser redpolls, that last listed as accidental. The Acanthis debate is certain to outlive us all.

 Familiar at least as a target bird to observers in Middle America, the old Prevost ground sparrow is no more. In its place, we have the white-faced ground sparrow and the Cabanis ground sparrow, the former occupying a range from southern Mexico to Honduras and the latter restricted to Costa Rica’s Central and Turrialba Valleys. The two species differ conspicuously in head and breast pattern — conspicuously, that is, if you’re fortunate enough to get a good look at these often sneaky sparrows.

And speaking, inevitably, of sparrows, the American birds going under that slippery English label are now assigned to a family of their own, PasserellidaeIn this, the AOS follows the recent practice of nearly all ornithologists over the past five years. It seems likely that the name will be replaced in the near future by Arremonidae, which if valid has nomenclatural priority.

Yellow-breasted Chat

The nine-primaried oscines — the “songbirds” at the back of the bird books — have also been rearranged, giving us all a new sequence to memorize. (I understand that the new sequence will be used in the seventh edition of the National Geographic guide, coming in a few weeks.) The most notable taxonomic change here is certainly the elevation of the yellow-breasted chat to its own family, Icteriidae, occupying a position in the linear sequence just before the orioles and blackbirds, Icteridae. This is just the latest stage on a classificatory journey sure to continue for a long, long time.

There will be more to say, no doubt, when the complete text of the supplement is readily available on line. Meanwhile, much to ponder.

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The Fifty-sixth AOU Supplement

It’s here, right on time, and birders around the world are scrutinizing every last densely printed word in this year’s supplement to the AOU Check-list. (That’s right: hyphen, small “l.”) There aren’t terribly many species-level splits or lumps this time, but there are some very significant changes at the higher levels of taxonomy, including some that will have many family listers spending the rest of the day on Travelocity.

Fully eighteen species are added to the North American list this time around, by the acceptance of new records or the taxonomic splitting of species already on the list. The “new” bird most likely to crest most birders’ horizons most immediately is the good old Egyptian goose, added here (following its addition to the ABA list last year) on the basis of established populations in Florida and California. Feral birds, park birds, and recent fugitives of this species can be seen anywhere in North America, though, and if the pattern seen in western Europe holds true, this clunky but oddly beautiful goose is poised to take over the continent.

Aiguamolls
Egyptian geese in Catalonia

There is tantalizing mention here of “a family-wide revision of English group names based on a complete phylogeny of the Trochilidae,” suggesting that sooner or later we may have to relinquish some of the evocative hummingbird names created by the French natural historians of the eighteenth and nineteenth centuries. For the moment, though, we get two hummingbird re-splits north of Panama. The former long-billed hermit now constitutes two species, the long-billed hermit (!) and the Mexican hermitPhaethornis mexicanus. These two ill-named birds differ “in vocalizations, behavior, genetics, and morphology” — and now, like many of us, I’ve got to dig up my notes and figure out which I’ve seen.

The other hummingbird split will be of particular interest to birders in The Bahamas and — get this — Pennsylvania. The woodstar on the islands of Great and Little Inagua is once again considered distinct from the widespread Bahama woodstar, and named, sensibly, the Inagua woodstarCalliphlox lyrura. All the US records are here suggested ex silentio to pertain to the Bahama woodstar in its new, strict sense.

It is worth pointing out that the citation the supplement gives for this woodstar’s original description is poorly formed; it should indicate that Gould published the name in the fourth volume of the fourth series of the Annals and Magazine, and that the name appears on pages 111-112. I expect that this will be corrected in the printed supplement.

In other trochilid news, the spelling of the steely-vented hummingbird‘s species epithet is corrected to saucerottei, with one “r,” which is how Nicolas Saucerotte and his family have always written it 

As if a Bahama woodstar weren’t enough, Pennsylvania birders also get to adjust their lists to take into account a seabird split. Pterodroma heraldica and Pterodroma arminjoniana are henceforth to be considered two species, the first known as the herald petrel in English, the second as the Trindade petrel. It is that latter tubenose that has occurred in the north Atlantic, and even inland in the eastern US, including, perhaps, Pennsylvania. Not only do we all need to learn how to spell “Trindade” now, but we’re going to have to come to a civil consensus about how to pronounce the island’s name.

Some of the big shifts this year are at the genus level. The lovely white-tailed hawk is no longer a Buteo but a Geranoaetus, a genus it shares with the variable hawk and, strikingly, with the black-chested buzzard-eagle. And in spite of their superficial similarity, the broad-winged hawk and the roadside hawk no longer nestle close to each other; the white-eyed southern bird is now in the genus Rupornis, where Kaup put it — naming Rupornis as a subgenus of his Asturina — more than 170 years ago.

Roadside Hawk Panama May 2007 288

The sequence of the big, blunt-winged hawks has been altered, too. North of Mexico, it now goes Harris’s, gray, red-shouldered, broad-winged, short-tailed, Swainson’s, zone-tailed, red-tailed, rough-legged, and ferruginous. To the delight of the makers of field checklists everywhere….

Ferruginous Hawk

A re-arrangement of the Hawaiian honeycreepers adds several new genera to the Check-listVestiaria is lost, but Akialoa, Chlorodrepanis, and Viridonia are given full rank, making for a total of 20 genera in Drepaninae. Several new honeycreeper species are recognized — eight of them, alas, extinct or likely so.

Closer to home for most of us, the American tree sparrow has been moved to its own monotypic genus, the blandly named (and hard to pronounce, I think) Spizelloides. This is the third name the species has had in the Check-list, from Spizella monticola to Spizella arborea and now to Spizelloides arborea. I’m not especially happy with the new species name: I suppose that the epithet could be argued to fall under the exception in ICZN 30.1.4.4., but the AOU treats all other generic names in -oides as masculine, and it should this one, too.

[Fix, July 7: I should have spent more time reading Klicka et al. and less time snuffling around in Greek dictionaries and the ICZN. In their paper, the authors explicitly state that Spizelloides is grammatically feminine, fully satisfying the 30.1.4.4. exception.]

American Tree Sparrow

The family of this and the other American sparrows — still, at this writing, Emberizidae, but likely to change soon, I suspect — is suddenly a lot smaller on the appearance of the new supplement. A long list of erstwhile sparrowish birds, all of them tropical, have moved from that family over to the Thraupidae. Yes, grassquits, seedeaters, the Caribbean bullfinches, the orangequit, the St. Lucia black finch, the Cocos and slaty finches, the peg-billed finch, the flowerpiercers, the yellow-finches, and the grass-finches are all true tanagers. The bananaquit, always among the bounciest of taxonomic balls, has also joined the tanagers.

Meanwhile, half a dozen other genera once placed among the tanagers have now taken uncertain seats, pending their elevation — each of them — to family status. Among these aspirants are the stripe-headed tanagers of the genus Spindalis, one species of which — the western spindalis — makes it into Florida. Family listers not holding all of these taxa in escrow will be doing some traveling soon.

Many birders did travel to see two cranes over the past several years. Neither, however, makes it onto the main AOU list. Both the California demoiselle crane of 2001-2002 and the hooded crane (or cranes) that wandered the continent between 2010 and 2012 have gone into the appendix — not, mark well, because the committee considers them definitely escaped captives, but because we do not know the origins of these birds. In this, the AOU echoes the ABA’s list, which treats all of these records as “questionable” based on provenance. But we most certainly have not heard the end of this one, as a cogent article in this month’s Birding shows.

Northern Harrier

Birders being birders, it’s those “rejections” — to use a word the AOU committee carefully avoids — that are likely to draw the most attention and spur the most debate. The committee also declined to separate the harriers, leaving the northern harrier and the hen harrier still to be considered conspecific. There will be less of a brouhaha about the failure to split the painted buntings or the Le Conte’s thrashers (the name of the thrasher is misspelled in the supplement), largely, I suspect, because most birders didn’t even know such had been proposed. The northern cardinal split into six species seemed a bit extreme from the start, but I’m glad the proposal was made: I’m once again hearing birders in the American Southwest speak unself-consciously of the superb cardinal, a fitting name if ever there was one.

Northern Cardinal superbus

 

There’s a great deal more, as there is every year when the supplement appears. You can read the whole thing here. Enjoy it!

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