Markedness

As birders learn to look closer, we more and more detect birds that somehow don’t fit the categories of the birdy books. Individual variation accounts for some of that, but many of the odd birds we encounter are, or at least seem to be, hybrids and intergrades, the products of matings more enthusiastic than accurate.

Some such hybrids are pretty obvious, like this apparent Mallard x Northern Pintail.

In many cases, though, the putative parent species are so similar to begin with, and their offspring so variable, that distinguishing between a “pure” individual and a hybrid can be a real challenge.

This gull at Clover Point this past weekend, for example, showed the mix of Western and Glaucous-winged Gull characters typical of our local “Puget Sound Gull”: a darkish mantle, blackish wingtips above, faintly marked wingtips below, and an orbital ring mixed yellow and red.

And this goose, hanging out in a Tucson park this winter, is probably a Ross’s Goose — or the long bill with a slight “grin patch” might suggest hybrid ancestry.

These are old stories and familiar, but lately I’ve been thinking about when birders look for (and find) hybrids. What it comes down to is markedness, or which potential parent taxon is deemed the default. This varies geographically, of course: in the east, a reddish Northern Flicker will be scrutinized, while over much of the west it’s the apparently yellow-shafted birds that draw special attention.

But I don’t think that markedness is always just about rarity and vagrancy. Even in those areas–and there are more and more of them as time goes on–where, say, Snow and Ross’s Geese are equally expected, it’s only the apparent Ross’s that are inspected for signs of hybridization: how often have you heard anyone cautiously report “an apparently pure Snow Goose,” even in places like southeast Arizona where both white geese are uncommonish? Nobody ever objects that a tentative Snow Goose’s bill is a bit short or the head a bit round or the plumage a bit white.

Even more strikingly, here in Vancouver I notice that birders (by which I mean mostly myself) readily pass over the slightly uncommon Myrtle Warblers, while subjecting the more abundant Audubon’s Warblers to a much more thorough examination. Adult male Audubon’s types with white in the throat or lightly marked wing coverts? Probable hybrids! But I simply tick the Myrtles; even when I linger over a particularly snazzy one (they are really very beautiful), I more often than not wind up moving on without checking it over at all as carefully as I do the yellow-throated birds. But surely there are hybrids that more closely resemble Myrtles than Audubon’s, aren’t there?

The lesson that I’ve drawn from these musings? Start to treat all the birds I see as “marked,” as potentially something different and weird–as worth looking at more closely than I already do. Who knows what I’ll find now?

Share

Hoover’s Warbler

The Yellow-rumped Warbler deluge shows no sign of receding, and Jericho Park is pretty much crawling with chipping, singing, flycatching Audubon’s and Myrtle Warblers again today.

It’s important–well, I think it’s important–to remember that both Audubon’s and Myrtle are polytypic; thus, it’s incorrect to speak of “the Audubon’s subspecies” or “the Myrtle subspecies” of Yellow-rumped Warbler, unless, of course, you’re using the word in the plural. The Myrtle Warblers we see here in Vancouver, the breeding race of northern British Columbia, are Dendroica coronata hooveri, differing in measurements and in some plumage characters from their eastern, nominate-race cousins.

This subspecies was described in 1899 by Richard C. McGregor, an adoptive Californian who would later become famous as the doyen of Philippine ornithology. He named his subsp. nov. after his college friend Theodore J. Hoover, collector of the type specimen and the older brother of Herbert.

In preparing his original description, McGregor also used specimens taken by Henry Ward Carriger, an early California oologist. I don’t know much about Carriger–fill me in if you do–but I was greatly impressed to read that as early as 1898, he had recognized the differences in the call notes of Audubon’s and Myrtle Warblers, a distinction that even today not all birders are aware of.

The Californians were out in front even then.

Share

Just Rumps

Vancouver’sJericho Park was covered with warblers this afternoon, but try as I might, I couldn’t find anything other than Yellow-rumps.

That’s an insidious “but,” of course, suggesting as it does that Yellow-rumped Warbler is somehow lesser than the other possible parulids. It is indeed much more abundant than any other, but that doesn’t make this species any less interesting. Or these species.

The birding world is abuzz with anticipation of a re-split of the yellow-rumped warblers, which were lumped almost 40 years ago in the great taxonomic massacre of the 1973 AOU Supplement. Now the Check-list Committee is evaluating a proposal to recognize at least two separate species in Dendroica coronata as now construed (and to correct the genus name to Setophaga–but that’s another story).

Both white-throated Myrtle Warblers and yellow-throated Audubon’s Warblers breed in British Columbia, and both are common migrants here in the Vancouver area, with the latter taxon generally the more abundant. Apparent hybrids and intergrades are easy to find if you look; one bird this afternoon had the yellow throat of an Audubon’s and the white malars and reduced white on the greater coverts of a Myrtle–I’m not sure I ever heard it chip, unfortunately.

The AOU Committee doesn’t much care what a birder wants, but still I hope for the split. And if it comes, I think we’ll all start paying attention again to what have for too long been “just yellow-rumps.”

Share

Golden Crowns and Black Tresses

Names are really just extreme words. And if the link between “ordinary” words and things is arbitrary, then that between names and the denoted can be downright capricious. Bird names are no exception, as generations of the literal-minded have moaned.

But a few birds enjoy names that are, wonder of wonders, straightforwardly descriptive.

Golden-crowned Sparrow? I’ll buy that. Hard to imagine what else you might call this bird with its, well, golden crown.

Unless, that it is, you happened to be Johann Friedrich Gmelin, who gave the species its scientific name in 1789. Gmelin was working from a not very good painting by John Latham, who labeled the bird “Black-crowned Bunting,” notwithstanding his description and depiction of the “fine yellow” of the crown.

Biodiversity Heritage Library

Gmelin followed Latham’s slightly misleading lead in assigning the species the epithet atricapilla, meaning “black hair.”

That too makes sense from some views, I suppose, though I can’t help wondering why Latham and then Gmelin would have zeroed in on those midnight locks rather than the aureate crown. No accounting for taste!

Share